World number one Williams on Saturday dominated Sweden's Johanna Larsson 6-2, 6-1 to surpass Martina Navratilova for most Grand Slam wins by a woman with 307.
Not only has she surpassed Navratilova, she matched Roger Federer's mark for men.
"To be up there with both men and women is something that's super-rare, and it actually feels good," said Williams, who said she was "really excited" to reach 307.
"Obviously I want to keep that number going higher," added Williams, who will get her chance when she takes on Kazakhstan's Yaroslava Shvedova for a quarter-final berth.
In the one hour it took to subdue Larsson, Williams appeared untroubled by the shoulder injury that has slowed her since her Wimbledon triumph.
"It definitely feels solid," she said. "I'm doing a lot of work on it so I can keep it in this position."
While Williams encountered little resistance as she moved a step closer to an Open Era record of 23 Grand Slam titles -- and Margaret Court's all-time record of 24 -- it was another story for the top men's seeds in action.
Wawrinka, a two-time Grand Slam winner and twice a semi-finalist in New York, had the closest call, saving a match point in a 4-6, 6-3, 6-7 (6/8), 7-6 (10/8), 6-2 victory over Britain's Dan Evans.
The 31-year-old third seed saved the match point at 5/6 in the fourth set tiebreaker, breaking the will of his 64th-ranked opponent.
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